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Copper activates endoplasmic reticulum stress The endoplasmic reticulum (ER) is essential for hepatocytes to maintain lipid metabolism, with its regulatory roles including: (1) ER membrane-localized sterol regulatory element-binding proteins (SREBP-1c, SREBP-2), which regulate DNL-SREBP-1c primarily mediates fatty acid synthesis, while SREBP-2 controls cholesterol synthesis

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This is achieved by recruitment of mTORC1 to the lysosomal membrane by GTP-bound RagA/B and GDP-bound RagC/D (activated by cytosolic amino acid sensors and lysosomal cholesterol sensors), and activation of mTORC1 on the lysosomal membrane by GTP-bound Rheb (activated by growth factors)